These differences shouldn’t affect most web developers, but if you need to know specifically, you can refer to the OS field of the User-Agent string. As Eric noted previously, the User-Agent strings will have the info you need:
IE7+ running on Windows Vista: Mozilla/4.0 (compatible; MSIE 7.0; Windows NT 6.0)
IE7 running on Windows XP: Mozilla/4.0 (compatible; MSIE 7.0; Windows NT 5.1)

Beta 2 is available now for developers and IT professionals who subscribe to MSDN or TechNet, as well as members of the Technology Adoption Program (TAP) and the TechBeta program. In the coming weeks, we’ll start the Windows Vista Customer Preview Program (CPP) for developers and IT professionals who aren’t part of one of those programs so they can get the code and begin their own testing. We’ll open up the CPP to a broader group of technology enthusiasts who’ll get a build of the OS to begin testing the various consumer scenarios Windows Vista enables.
Source: Microsoft Announces IE7+ for Vista
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